In Eastern Dan (< Southern Mande < Mande < Niger-Congo), the common Mande strategies of marking genitival relations are available, namely head-marking by means of tone, and dependent-marking (by a specialized postposition) in the alienable possession construction. However, against this common background, some important modifications have evolved. The opposition between the head-marked and the head-unmarked constructions, although retained, has become semantically blurred: in the proto-language, the head-marked genitival construction had the semantics of “modification-by-noun”, while in the Eastern Dan, this meaning has undergone erosion. In the dependent-marked constructions, a case-like opposition has emerged in the alienable possession constructions: different possessive markers (postpositions) are used depending on whether the head noun of an NP containing the possessive construction stands in the locative case or not. Other postpositions can also serve as markers of genitival relations. In addition, morphological case can sometimes serve to mark genitival relations.